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Knuckles is the partner of the Whistling Skull. He is a strong man with a learning impairment. His father provides him with special lotion to harden his skin, making it more resilient than normal human skin. 
His father was the Sixth Whistling Skull.
- 6 Appearances of Nigel Singleton (Earth-40)
- 5 Images featuring Nigel Singleton (Earth-40)
- 1 Quotations by or about Nigel Singleton (Earth-40)
- Character Gallery: Nigel Singleton (Earth-40)